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Average height of top tiller (Plant height)

Plant height was depressed in Mn deficient plants and removal of stress increased it significantly (Table 3). The maximum height was obtained in taller cultivars HD 2009 and C 306. Thus potential tallness appeared to be related to Mn stress tolerance in wheat. Effect of Mn was more pronouned on the growth of the top tiller in comparison to younger tillers. However in different Mn rates, the differences were insignificant except in highly susceptible cultivars DWL 5023 and KSML 3 which had significant increase even with 20 mg Mh kg-1. The plant height was positively related to grain and straw yields (r = 0.84** and 0.80** respectively).

The height of tillers, likely to contiribute towards increased assimilate production for grain, seemed to have acted as a secondary sink for assimilates, contributing more towards straw yields.

Sink capacity

If the assimilates are not limiting the sink capacity determines the ultimate grain yield. The sink capacity comprised of the interaction of the following factors:

Number of spikes (ears)

The spike number (number of fertile tillers) was very low in control plants of DWL 5023 - the only spike produced was devoid of grains (Table 4). The spike number was poorly related to straw and grain yields and thus did not serve as a good indicator of grain yield as the size of the spike and grain number in each differed widely.

Average spike length

Under Mn stress the spike length was very small in DWL 5023 but increased significantly with 5 mg Mn/kg application, but the differences between Mn rates were non-significant (Table 4). The grain yield was poorly related to spike length in control, implying possibilities of both impaired grain development as well as grain filling. However, in 5 and 10 mg Mn kg-1 supplied plants the grain yield was related to average spike length.

Number of grains in the top spike (Number of flowers fertilized)

The average number of grains was very significantly decreased with Mn stress, and these strikingly and significantly increased when Mn stress was removed (Table 5). All the cultivars produced maximum number of grains at 10 mg kg-1 Mn rate of application except DWL 5023, KSML 3 and C 306 which gave maximum grain number at 20 mg kg-1 Mn rate. The average grain number was highest in cultivar WG 377 followed by WL 711, WL 410, HD 2009, TL 419, WL 1562, C 306, KSML 3, WG 357 and DWL 5023 in decreasing order.

The highly susceptible DWL 5023 and moderately susceptible WL 1562 did not produce any grains under Mn stress. Failure to set grains in these cultivars could be ascribed to a single or a combination of the following factors:

a) Interruption of sink capacity

b) Improper translocation mechanism

c) Scarcity of leaf photosynthates

d) Infertile pollen/nondehiscence of anthers

e) Non-receptive stigmas

f) Absence of fertilization

g) Non-viability of embryos

h) Deficiency of growth regulators/ accumulation of inhibitors of cell division



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