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Long time breeding practice has revealed that transgressive inheritance in scab resistance is of frequent occurrence, but the so-called "resistance" is actually varying degrees of tolerance to the extension of infection and not infection itself. Owing to lack of source of resistance to infection (note: recently Agropyron ciliare and A. repens were found to be highly resistant to scab infection by Nanjing Agric. Univ. and Sichuan Agric. Univ., independently) and a certain extent of negative corelation between scab resistance and good agronomic characters, it is difficult to achieve even a small breakthrough in the improvement of scab resistance by the conventional method. Therefore recurrent selection by using Taigu dominant male sterile gene Ms2 or Ta1 has been adopted in recent years for pyramiding the major and minor genes which control scab resistance. Such a project was initiated and undertaken by Nanjing Agric. Univ. and its partners. They crossed 15 selected scab resistant varieties including landraces with five improved varieties carrying Ms2 gene to make a composite of 75 combinations. After three years of intermating a CO population was established which was grown in four locations under the same design. Phenotypic selection with an intensity of 5-10% was practised aiming first at scab resistance and paying considerable attention to major agronomic traits as well. Seeds from the selected male sterile plants were bulked with equal quantity to form the stock material for the next cycle. After three cycles of phenotypic selection comparison among the four populations from CO to C3 was made in Nanjing. The results showed that percent of infected plants in the population and the number of infected spikelets of infected plant reduded materially as the cycle advanced, with an average of 20% decrease per cycle for the latter. After two to three cycles of selection scab resistance of the population almost approached the level of Sumai 3, and spike characters except kernel weight were correspondingly improved with a gain per cycle varying from 2-8% (Jiang et al, 1992). Meanwhile, South China Agric. Univ. also conducted a similar study by crossing eight resistant parental stocks with some high yielding varieties carrying Ms2 gene. After several cycles of selection some promising lines designated as T474, T229 and the like with acceptable scab resistance and desirable agronomic characters were developed.


Breeding for semidwarfness

Early in 1957 Xianyang Prefectural Agric. Res. Inst. using Suwon 85 (bearing Rht1 and Rht2) as a source of dwarfhess developed the first series of semidwarf intermediate materials among which Xiannong 39 was a good representative. Then Northwest Agric. Univ. by crossing Xiannong 39 again with a promising hybrid line produced the first series of semidwarf varieties in which Aifeng 3 was the best one and released in 1971. Since then breeders from Shandong, Sichuan, and Henan provinces had developed in succession a number of semidwarfs which were widely grown in some major wheat regions. Those varieties listed hereinbefore for the Yellow and Huai Rivers valleys and Sichuan basin are all semidwarfs. Presently about 40% of the total wheat acreage is sown with semidwarfs.

In breeding for semidwarfness development of desirable intermediate or link-up materials is very important. In addition to Xiannong 39, there was another example. Shandong Agric. Univ. was successful in developing an intermediate, link-up material called Aimengniu which was a product of Aifeng 3//Mengxian 201/Neuzucht. It combined dwarfness from Aifeng 3 and multiple disease resistance from Neuzucht, but a little bit late in maturity. Subsequently breeders in the University using this as a source of dwarfhess and disease resistance in crossing bred a number of improved varieties known as Lumai series which were widely grown in Shandong province. Another experience in the breeding for semidwarfness is that selected crosses between short or midstatured varieties may produce some semidwarfs through transgressive segregation. For example, Huixian Red (short)/Abbondanza (medium) -> Taishan 4 (semidwarf); Youbao (short)/036 (tall, a Beijing selection) -> semidwarf transgressive segregantsGand others.

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