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Allelic variation has been demonstrated at all of the Gli-2 loci, including 13 alleles at Gli-A2, 11 at Gli-B2, and 10 at Gli-D2 in a study of 39 varieties(657).



The location of Gli-R2 in S. cereale in thought to have envolved from S. montanum (659) via a translocation between 2R and 6R(663).



2. Other

Three other prolamin loci have been identified on the short arms of the group 1 chromosomes of hexaploid wheat but the relationship between them is not clear. A gliadin locus, designated Gld-B6, was described in CS/Thatcher substitution lines and mapped 28 mu proximal to Gli-B1 (593). Another locus, designated Gld2-1A (760), controlling production of proteins was described in a cross between Bezostaya 1 and Gernot and mapped 31 recombination units from Gli-A1 (664). A glutenin locus in a similar position on 1BS, 22 mu proximal to Gli-B1, was designated as Glu-B2 and mapped in a Sicco x Chinese Spring cross (599).

A set of two wheat endosperm storage gene loci, Tri-A and Tri-D1 (762), were located 11.0 + or - 1.8% and 10.1 + or - 2.2% from the centromeres on 1AS and 1DS, respectively (666). Allelic variation occurred at both loci(762).


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