The complexity of the electrophoretic patterns of the products of these loci currently defies the naming of alleles. Nevertheless, variation has been described (644, 657) and applied in mapping experiments (646,320,60,593,649). Sixteen combinations of Gli-B1 and 4 combinations of Gli-D1 subunits are listed in 641. Multiple alleles are described in 657, 15 at Gli-A1, 18 at Gli-B1, and 8 at Gli-D1. An indication that the gliadin genes and the LMW glutenin genes, at the loci presently described as Gli-1, may be separable was provided by the report of recombination (1.7%) within Gli-B1 (666).
In barley, the B and C hordeins are controlled by the Hor 2 and Hor 1 loci, respectively, which are linked (656) on chromosome IHS (652, 565). The map distances and homology of the proteins indicate that Hor 1, the locus closest to the centromere, is partly equivalent to Gli-1 (526).
