National BioResource Project (NBRP)::KOMUGI -Integrated Wheat Science Database- : Genome / Development

Gene symbol	Class				Synonyms	Chromosomes	Germplasms
Gene symbol	Level1	Level2	Level3	Level4	Synonyms	Chromosomes	Germplasms
Q	Gross Morphology: Spike characteristics	Squarehead/spelt			k	5AL	bin:5AL-17,v:Common wheats. CS,v:Iranian spelts,tv:T. turgidum ssp. carthlicum, durum and polonicum,ma:Complete linkage with cDNA clone PtAq22,ma:Q was cloned and shown to have similarity to AtAP2 (APETALA 2) ,the Q allele was more abundantly transcribed than the q allele transcription factors
Q	Comment:
q	Gross Morphology: Spike characteristics	Squarehead/spelt			K		v:Macha wheats,v:European spelt wheats,v:vavilovi wheats,s:CS^8/White Spring Spelt 5A,tv:T. turgidum* ssp. dicoccum, dicoccoides,ma:Cent - Xrsq805(Empb)-5A - 4.6 cM - Q - 4.3 cM - Xpsr370-5A,ma:Q was physically mapped in 5AL, fraction length 0.87, bracketed by deletions 5AL-7 and 5AL-23,ma:Q - 9.3 cM - Xpsr370-5A
q	Comment: The speltoid phenotype of at least some spelts may be caused by genes at other loci {0140}. Fine mapping of the 20 cM region possessing Q and delimited by deletions 5AL -7 and -23 is reported in {0324}.
C	Gross Morphology: Spike characteristics	Club/Compact spike			Cd	2D,2DL,probably 2DL	bin:C-2DS1 - C-2DL3, markers flanking C were located on either side of the centromere,i:S-615^11/Elgin,s:CS^6/Poso 2D,s:CS^5/Red Egyptian 2D,v:Club wheats,v:Coda,v:Corrigin,ma:Coda / Brundage: Xwmc144-2D* - 1 cM - C - 8 cM - Xwmc18-2D,ma:Corrigin / CS (tauschii 2D): Xwmc245-2D - 1 cM - Xcfd16-2D / Xgwm358-2D / C / Xcfd-2D - 1 cM - Xbarc145-2D
C	Comment:
s2	Gross Morphology: Spike characteristics	Sphaerococcum			sp2		v:Sphaerococcoid wheats. "Sphaerococcum simulator"
s2	Comment: Partially dominant{1286}. Sphaerococcum-like tetraploid wheats were reported{122,475,1282,1286}, but comparisons between them, or with s2, were not made. Whereas Schmidt & Johnson{1281} reported a single recessive controlling the sphaerococcum character in tetraploid wheat, Joppa{621} using the same stock found that two recessive genes were necessary to produce this phenotype.
bh-A1	Gross Morphology: Spike characteristics	Branched spike			bh	2AS	tv:PI 349056
bh-A1	Comment:
P1	Gross Morphology: Spike characteristics	Elongated glume			P,Eg,P-A^pol1,P-A^pet1	7AL,7A or 7B (based on linkage of 0.2 with a gene for red coleoptile)	i:Saratovskaya29^8//Novsibirskaya 67^2/T. polonicum,itv:P-LD222 = LD222^11/T. turgidum* var polonicum,tv:T. polonicum,tv:T. petropavlovskyi,ma:Xgwm260 - 7A(S) - 2.3 cM - P1 - 5.6 cM - Xgwm1083-7A(L),ma:Xgwm890 -7A - 2.1 cM - P1,ma:Xgwm260-7AS - 2.3 cM - P1^pol - 5.6 cM - Xgwm1083-7AL,ma:Xgwm890-7AS - 2.1 cM - P1^pet
P1	Comment: Note: The loci determining elongated glumes in T. turanicum and T. durum conv. falcatum are not homoeologous to the P loci in the centromeric region of the group 7 chromosomes{0254}.
P2	Gross Morphology: Spike characteristics	Elongated glume				7BL	itv:LD222^7/T. ispahanicum,tv:T. ispahanicum*
P2	Comment: According to {0254} the loci of T. polonicum, T. petropavlovsky and T. isphanicum are allelic ('homoeoallelic') whereas other workers had claimed genes in the first two forms were not allelic. Wang et al. {0254} however concluded that loci bearing alleles for elongated glumes in T. turanicum and T. durum conv. falcatum were not part of the above series.
S-A1	Gross Morphology: Spike characteristics	Sphaerococcum				3A	v:CS
S-A1	Comment:
S-A1a	Gross Morphology: Spike characteristics	Sphaerococcum					v:CS,v:common wheats
S-A1a	Comment:
S-A1b	Gross Morphology: Spike characteristics	Sphaerococcum			S3		v:MS 1453,ma:Xgwm2-3A(S) - 5.1 cM - S-A1 - 6.6 cM - Xgwm720-3A(L)
S-A1b	Comment:
S-B1	Gross Morphology: Spike characteristics	Sphaerococcum				3B	v:CS
S-B1	Comment:
S-B1a	Gross Morphology: Spike characteristics	Sphaerococcum					v:CS,v:common wheats
S-B1a	Comment:
S-B1b	Gross Morphology: Spike characteristics	Sphaerococcum			S2,s¹⁶²¹⁹		v:MSK 2452,v:MSK 2454,tv:MA 16219,ma:Xgwm685-3B(S) - 4.2 cM - S-B1 - 0.5 cM - Xgwm566/Xgwm845/cent
S-B1b	Comment:
S-D1	Gross Morphology: Spike characteristics	Sphaerococcum				3D,3DS,3DL	v:CS
S-D1	Comment:
S-D1a	Gross Morphology: Spike characteristics	Sphaerococcum					v:CS,v:common wheats
S-D1a	Comment:
S-D1b	Gross Morphology: Spike characteristics	Sphaerococcum			s1, sp1		i:S-615^11/T. sphaerococcum* var. rotundatum,s:CS^7/T. sphaerococcum rubiginosum* 3D,v:Sphaerococcum wheats,v:T. antiquorum K056397 & K56398
S-D1b	Comment:
S-D1c	Gross Morphology: Spike characteristics	Sphaerococcum			S1		v:MS 3287,ma:Xgdm72-3D(S) - 8.0 cM - S-D1 - 2.9 cM - Xgwm456-3D/cent
S-D1c	Comment:
QEl.ocs-5A.1	Gross Morphology: Spike characteristics	Ear length				5AL	v:CS(T. spelta 5A)/CS(Cappelle-Desprez 5A) RI mapping population,ma:Associated with Xbcd9-5A
QEl.ocs-5A.1	Comment:
bh-D1	Gross Morphology: Spike characteristics	Branched spike			mrs	2DS	bin:2DS-5 0.47-1.0,v:Ra1,v:Ruc163-1-02 = Ra1 / ZGK242-81,v:RUC163167-1-02 = Alana /3/Ra1/ZGK242-82 // Ra1,ma:Xwmc453-2D / bh-D1 - 7.8 cM - Xgwm988-2D,ma:Xwwm484-2D - 3.3 cM - Xwmc453-2D/bh-D1 - 3 cM - Xwgm988-2D
bh-D1	Comment: Ra1 is a mutant stock maintained at the NI Vavilov Research Institute of Plant Industry, St Petersburg, Russia. A chromosome 2B gene of minor effect was identified {9907} . In a monosomic analysis of the hexaploid line LYB with supernumerary spikelets, Peng et al. {9908} located recessive genes in chromosomes 2A and 4A that promote the development of supernumerary spikelets and a gene in chromosome 2D that prevents their expression.
bh-R1	Gross Morphology: Spike characteristics	Branched spike			mo	2R	al:S. cereale D40,ma:Xrms056-2R - 15.7 cM - bh-R1 - 10.7 cM - Xcfe209-2R
bh-R1	Comment:
Pis1	Gross Morphology: Spike characteristics	Multi-gynoecium				2DL	bin:C-2DL3-0.49,v:TP Mutant,ma:Xgwm539-2D - 17.6 cM - Pis1 - 19.5 cM Xgwm349-2D
Pis1	Comment:
C_g	Gross Morphology: Spike characteristics	Club/Compact spike				2BL	bin:2BL-0.48-0.89, near breakpoint 0.69,v:Akage Gumbai,v:Akage Gumbai 22,v:Gumbai 22,v:Kinoshita Komugi,v:Nakote Gumbai,ma:Xhbg410/Xhbg440-2B - 18.1 cM - C_g - 15.3 cM - Xgwm47-2B
C_g	Comment: