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Materials and methods

The plant materials used in this investigation are listed in Table 1. To transfer leaf rust and stripe rust resistance from diploid Aegilops species with C or U genomes to hexaploid wheat, amphiploids were developed between these species and susceptible T. durum cultivars. To synthesize amphiploids, the coleoptiles of 4-5 day-old F1seedlings, of these crosses, were treated with 0.25% colchicine in 2% DMSO solution for four hours (Gill et al. 1988).These amphiploids were used as the bridge for the transfer of leaf and stripe rust resistance genes into cultivated hexaploid wheat. To induce homoeologous chromosome pairing these amphiploids were first crossed with T. aestivum cv. Chinese Spring carrying the PhI gene from Ae. speltoides (Chen et al. 1994; Aghaee-Sarbarzeh et al. 2000) and further backcrossed to Chinese Spring.

The parents, F1's, amphiploids, and the derivatives of their crosses with Chinese Spring (CS) were scored for field reaction to leaf rust and stripe rust by recording terminal disease severity and response to individual rust pathotypes. The disease severity under field conditions was recorded as percentage of leaf area covered by rust following modified Cobb's scale as developed by Peterson et al. (1948). According to this scale, at 100 % disease severity, the actual leaf area covered by rust pustules is 33 %. The response of individual plant/entry was recorded as follows; 0: resistant, no infection, R: resistant, necrotic areas with or without minute uredia, MR: moderately resistant, small uredia surrounded by necrotic areas, MS: moderately susceptible, medium sized uredia with no chlorosis, S: susceptible, large uredia without necrosis or chlorosis, X: intermediate, variable size uredia, some with necrosis or chlorosis. In addition, these materials were also evaluated for seedling resistance to individual pathotypes of leaf rust and stripe rust. First leaves of five to seven day-old seedlings were inCulated with urediospores mixed with talc using lancet needle (Nayar et al. 1997). The inCulated seedlings were incubated for 24 hours at 20 plus or minus 1C for leaf rust and 8-9C for stripe rust at 100 % relative humidity. Subsequently, plants were placed on benches in the glass house at temperature around 20-25C for leaf rust and 15-20C for stripe rust. About 11 to 15 days after inCulation, infection types were recorded according to a modification of 0-4 scale (Knott 1989) given by Stakman et al. (1962) as follows; 0: immune, no visible infection, 0;: no uredia, hypersensitive flecks present, 1: resistant, minute nonsporulating uredia surrounded by distinct necrotic areas, 2: resistant to moderately resistant, small uredia with slight sporulation surrounded by chlorotic or necrotic areas, 3: moderately resistant to moderately susceptible, medium sized sporulating uredia, chlorotic areas may be present, 4: susceptible, large sporulating uredia with no chlorosis or necrosis, X: resistant, heterogeneous, variable size uredia distributed over leaves. Variations were indicated by the use of - (less than the average for the class) and + (more than the average for the class) as superscripts.


Results and discussion

The accession 13749 of Ae. umbellulata was resistant under field conditions as well as at seedling stage to pathotype N of stripe rust and pathotypes 77-4, 77-5, 104B, and 104-2 of leaf rust (Table 2). However, the durum wheat cultivars, Bijaga Yellow (BY) and Malvi LCal (ML) were susceptible to both the rusts under the field conditions. The cultivar BY was resistant to races 77-4 and 77-5 of led rust and race N of stripe rust, but susceptible to races 104B and 104-2 of leaf rust, whereas, the durum wheat cv. ML, showed susceptibility to all the pathotypes of rusts , except 104B of leaf rust, at seedling stage (Table 2).

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