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Materials and methods

Initially, six cultivars (WC 29, WH 291, SGP 14, RAJ. 1972, WH 377 and HD 2329) of spring wheat were crossed in all possible combinations, excluding reciprocals, and their
15 F1's were grown in a Randomized Block Design with three replications. Five plants in each replication were selected for recording observations. Six basic generations (P1, P2, F1, F2, BC1, BC2) of three crosses viz, WC 29 x WH 291, SGP 14 x RAJ. 1972 and WH 377 x HD 2329 selected on the basis of combining ability were developed to determine gene effects and to predict the frequencies of transgressive segregants. The six basic generations were grown in a Randomized Block Design with three replications. Five plants in each non-segregating generation, 25 in each backcross generation and 50 plants in each F2 population were taken for recording observations. To predict frequency of transgressive segregants using F3 family data, a random sample of 30 F2 plants was taken to produce 30 F3 families of each cross. The parents and progenies were grown in an Augmented Design with six blocks. Fifteen plants in each F3 families of each cross and five plants in each of the parents in each block were taken at random for recording observations.

To identify transgressive segregants in the F2 population, the parents, F1's and F2 population of each cross were grown in a Randomized Block Design with three replications. Five plants in each of the non-segregating generations and 350 plants in each of the F2 population in each replication were selected randomly for recording observations. The F3 progeny of selected transgressive segregants were grown for progeny testing.

To identify transgressive segregants in biparental progenies (BIPs) and F4 bulks, the parents, BIPs and F4 of each cross were grown in an Augmented Design with six blocks. Five plants of each parent in each block and 450 plants in each population of each cross were taken at random to identify transgressive segregants.

In each experiment, the row length was 6m, spaced 25 cm apart. The plant to plant distance within rows was 15 cm. To ensure leaf rust development, spreader rows were planted around each replication in each experiment. Spreader rows were inoculated artificially with a mixture of races of leaf rust of wheat. The observation on leaf rust was recorded following Loegering (1959). Field resistance expressed as a coefficient of infection was computed as described by Yadav (1984).

Analysis of variance for Randomized Block Design and Augmented Design was done following Addelman (1969) and Federer (1961), respectively. Combining ability analysis was carried out following Method 4, Model 1 of Griffing (1956). Initially, generation mean analysis following three parameter model as suggested by Cavalli (1952) was carried out. A six parameter model suggested by Hayman (1958) was applied if the three parameter model was found to be inadequate.
The frequency of transgressive segregants was predicted from generation mean analysis and F3 family alone following Jinks and Pooni (1976, 1980).

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