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Wheat Information Service
Number 72: 18-24 (1991)


Ditelosomic analysis of some morphological characters in wheat, Triticum aestivum cv. 'Chinese Spring'

Afshan Afzal and Ahsan A. Vahidy

Department of Genetics, University of Karachi, Karachi- 75270, Pakistan


Introduction

Aneuploidy, which is the loss or increase of chromosomes has been widely studied in hexaploid wheat, Triticum aestivum (2n = 6x = 42). Aneuploids can be classified as primary or secondary, the former being a loss or increase of complete chromosomes, the latter involve different dosages of chromosomes derived through the misdivision of the centromere.

Due to the availability of the aneuploid series in 'Chinese Spring', which was developed by Sears (1954), the complexity of genetic analysis in hexaploid wheat has been greatly reduced. All the 42 possible telocentrics have been obtained (Sears and Sears 1978). Out of these, 35 are maintained as ditelosomics. They have been obtained by selfing either monotelosomics or monotelodisomics (Sears and Sears 1978). Ditelosomics, in which each line is deficient in one pair of chromosome arms, breed true and are usually of high fertility (Law et al 1987). These are most frequently used to study the role of individual chromosomes, as differences for a particular character can be directly attributed to the effect of the missing arm.

Days to heading, whose nature of dominance is not clear yet, is a complex character. Date of sowing, day length, latitude and day and night temperatures are some of the factors affecting days to heading. Spike length is a genic character. Genes controlling spike length have an additive effect. Leaf size has a direct relationship with biomass and yield of the plant. As compared to other leaves, the flag-leaf makes an important contribution of photosynthates especially at the stage of grain-filling. Hence elucidation of the role various chromosomes play in determining flag-leaf size is important.

In this paper, the contribution of chromosome arms in the performance of some agronomic characters, viz., days to heading, spike length, flag-leaf length and width was estimated by comparing ditelosomics with normal disomics in T. aestivum cv. 'Chinese Spring'.


Materials and Methods

The material for the present study comprised of 32 ditelosomics, which could be maintained and confirmed cytologically, and disomics of T. aestivum cv. 'Chinese Spring'. The seeds of the ditelosomics were kindly provided by Dr. E. R. Sears, University of Missouri, Columbia, U.S.A., and maintained at the Department of Genetics, University of Karachi, Karachi, Pakistan.

The seeds were placed on moist filter papers in Petri dishes and kept at 20C. Root tips were collected after 3 days for cytological confirmation. The seeds were resown in small pots and kept at 20C. After 15 days, the plants were transferred to large pots and kept in the screen house, i.e. in the open, under natural conditions.

The day on which the spike emerged from the boot leaf was taken as date of heading from which days to heading was calculated. Observations on spike length, and flag-leaf length and width were recorded at maturity. Mean and standard error for these variables in each line were calculated. The statistical analysis was carried out using the statistical package SPSS/PC+ and utilizing the command MANOVA. The analysis of variance was done by using the Completely Randomized Design (CRD). Several types of contrasts are available for different types of parameter estimates. The contrast used in our analysis was 'SIMPLE' in which one variable (in this case the disomic) was compared to the rest (the ditelosomics). This type of analysis is not orthogonal (Norusis 1988).

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