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Results and Discussion

Experimental results revealed that germination rate in all the varieties showed a slight increase at lower doses (1 - 3 kR) as compared to the control, whereas it decreased at higher doses (
Table 1). More reduction was seen in durum wheats.

Seedling height is often used as a measure of radiation damage in seeds. In all the varieties there was a stimulation in seedling growth at lower doses (1 - 3 kR) as compared to the control, whereas seedling height decreased as the dose increased from 5 kR to 30 kR (
Table 1). Similar results were observed for mitotic index.

Stimulation in germination may be due to the formation/activation of some stimulatory substances e.g. auxins, which take active part in germination (Simonis 1966), whereas the reduction may be caused by radiation induced damage through the alteration in endogenous hormonal levels (Mishra et al 1980). Some of the damage could also be due to a direct effect of radiations in killing meristematic cells. Stimulation in seedling height may be due to faster cell division, change in auxin balance or enhancement in enzyme activity. Radiations are capable of considerably changing the type of information of certain DNA molecules (Simonis 1966). This may be the reason for the increase in mitotic index at lower doses and a decrease at higher doses.

Various types of abnormalities observed during mitosis in somatic cells of root tip were in the form of laggards, bridges and chromatin stickiness (
Table 2). These increased with an increase in radiation dose. The most frequent abnormality was in the form of laggards at anaphase, and the second was bridges. During meiosis cells with univalents at metaphase was observed most frequently, and the second was the cells with laggards at anaphase (Table 3). Univalents may result either from the failure of chromosomes pairing at zygotene (asynapsis) or from the disjunction of homologous chromosomes at diplotene (desynapsis), because chiasma formation did not occur. Possible explanation for the formation of bridges may be that broken ends containing centromere from two different chromosomes unite to form dicentric chromosome, or that chromosomes are clumped together due to stickiness and are often unable to separate compelety at anaphase. Chromatin stickiness may be caused by some changes in the surface properties of the chromosomes which caused them to adhere to each other on coming in contact (Khanna 1986).

Considering the overall data, it appears that hexaploid wheats seem to be a little more radiosensitive as compared to the tetraploid wheats. This could be due to a greater chromosomal volume in the hexaploid Wheat. Gupta and Roy (1985) reported that gamma-rays induced more meiotic abnormalities in tetraploid Physalis peruviane as compared to the diploid P. ixocarpa.

References

Gupta SK and Roy SK (1985) Comparison of meiotic abnormalities induced by gamma rays between a diploid and tetraploid species of Physalis. Cytologia 50: 167-175.

Khanna VK (1986) A comparison of radiosensitivity of wheat and triticale. Acta Bot Indica 14: 110-114.

Mishra SD, Mathew T, Joshi RK and Gaur BK (1980) Radiation induced delay in organogenesis of Kalanchoe daigremontiana leaves. Env and Exptl Bot 20: 213-215.

Simonis W (1966) Physiological problems related to the effects of small doses of radiation on plants. In: Effects of low doses of radiation in crop plants, IAEA, Vienna.

Swaminathan (1964)

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