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Spike length in mutant strains derived from different cultivars significantly enhanced as compared to their respective controls, except mutant strain No. 10 of Nayab in which spike length is signfiicantly (P>=0.05) reduced whereas, in mutant No. 13 of Indus-66 spike length was non-significantly decreased. Mean spike length of mutant strains derived from C-591 cultivar were reasonably homogenoeous. Similar significant shifts in mean values for spike length was also observed by LARIK (1975a, b,. SIDDIQUI and GHAFOOR (1974).

For spike length, the mutant strains derived from mutagenic treatments of Indus-66 cultivar exhibited an increase of genetic variance as compared to all other mutant strains, this could be due to low yielding mutant strains (Table 2). Similar conclusions have also been drawn by GAUL et al. (1966) and SIDDIQUI and GHAFOOR ARAIN (1974). The heritability and genetic gain expected from selection also showed same tendency as the genetic variance. These estimates were of higher magnitude for the mutant strains of Indus-66 cultivar compared with C-591 and Nayab control. Thus indicated greater gains from selection for spike length are anticipated among the strains of Indus-66 (GHAFOOR ARAIN, 1973).

Spikelets per spike, seeds per spike and seed index (1000-seed wt.) are important yield components and are considered a reliable measure of yielding ability BOROJEVIC and BOROJEVIC (1972) as the frequency of induced changes in next generation depends on the number of seeds which transmit them (LARIK, 1978). The best criteria for the evaluation of superior genotypes is a comparison of its 1000-seed weight with the mother variety (GUSTAFFSON et al, 1971). Mean values for these quantitative characters were significantly (P>=0.01 and 0.05) reduced, except mutant strain No. 44 and 37 of Nayab and Indus-66 for spikelets per spike, mutant No. 44 and 13 of Nayab and Indus-66 for seeds per spike and mutants No. 27 and 44 of Nayab for seed index. These mutant strains displayed significant (P>=0.01, 0.05) increase for these quantitative characters over their respective controls. In the present study, the shift of mean values for these quantitative characters is mostly in negative direction, it provides support for the hypothesis of GAUL and AASTVEIT (1966) which states that the change in the mean values for the quantitative characters occurs both in positive and negative direction and is associated with reduced vitality independent of the genotype used.

Spike length of mutant-13 of Indus-66 and seeds per spike of mutant-7 and 28 and mutant-27 of C-591 and Nayab respectively were not significantly alterd from that of their respective controls. The non-significant shift in mean values of these characters is in agreement with findings of GHAFOOR ARAIN and SHEPHERD (1977) in wheat and OKA et al. (1958) in rice.

Highest genotypic co-efficient of variation was given by mutant strains No. 37 of Indus-66 for spikelets per spike, seeds per spike and seed index, indicates the potential for the improvement of these characters in the mutant populations. Heritability and genetic advance were of higher magnitude for the mutant strain No. 37 of Indus-66 for spikelets per spike and seed index and mutant No. 44 of Nayab for seeds per spike. This indicates that greater gains from selection for these characters are anticipated among the strains of Indus- 66 and Nayab.

The present studies conclude that selection for superior genotypes for yield and yield components having low heritability values and low genetic advance is very difficult. It is therefore imperative from the breeders point of view to select those attributes displaying high heritability values among the mutant strains. The present studies have provided an evidence on the induction of genetic variability connected with yield and yield components of wheat crop. Thus, induced genetic variability can effectively be exploited for evolving mutant strains possessing desirable attributes.



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