| Amphidiploid from the hybrid, Aegilops umbellulata
4x x Aegilops squarrosa 4x Masatake TANAKA Laboratory of Genetics, Faculty of Agriculture, Kyoto University, Kyoto, Japan All the possible amphidiploid combinations between five groups of principal analyzers (C, Cu, D, M and S), except Cu x D, have been already obtained (WIS No. 1). M and Mu are included in M group, and S, Sb and Sl in S group. In 1956, I synthesized an amphidiploid CuCuDD from the cross of induced 4x parents, Aegilops umbellulata 4x x Ae. squarrosa 4x. But, in this amphidiploid the chromosome pairing was not regular, and often univalents and polyvalents were seen. The seed-fertility was rather poor (ca. 13%) From the morphological point of view, this amphidiploid bears the characteristics of Section Vertebrata, resembling Ae. juvenalis. Consequently, CuCuDD x M group (Ae. comosa, Ae. Heldreichii and Ae. uniaristata) must give rise to an amphidiploid which corresponds to Ae. juvenalis, as Kihara (1954) indicated. |