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These uniform Iranian varieties have also become of great interest to breeders in other countries including in Europe. According to the investigations of Mrs. Dr. E. FUCHS, Biologische Bundesanstalt, Braunschweig, West-Germany (information through correspondence), the 4 varieties proved to be susceptible to race 20 Iran of Puccinia strii formis but resistant to race 20 Turkey ; 2 varieties are resistant to 3 races prevalent in Europe.

Table 3. Seedling tests of 4 Iranian wheat varieties with different races of yellow rust

In 1952 growing of Triticum aestivum ssp. spelta was found on the high plateau of Shahr- Kord near Isfahan. The extent of this spelta growing area was determined by taking samples from 18 localities which are pintpointed in the map by dark circles (Fig. 1). The genetic variation of these samples was analysed in a research programme as to the variation of the spelta-complex and other characters such as winter-summer type etc. The spelta complex is some times broken up, and great numbers of transitorial types to the free threshing T. aestivum ssp. vulgare could be detected. Thus a contribution could be given on the problem-being yet unsolved at that time-of the evolution of the hexaploid wheat as to whether T. aestivum ssp. spelta ist the ancestor of the free threshing hexaploid wheats or a derivative which has arisen from free threshing types by mutation recently occurred in Europe. Since the analysis of the spelta findings in Iran had been made the latter hypothesis forwarded by SCHIEMANN and FLAKSBERGER simultaneously in 1930 is now to be abolished (GOKGOL 1961, KUCKUCK 1964, KUCKUCK und SCHIEMANN 1957, POHLENDT 1958). Spelta growing in Iran has disappeared completely in the course of the past 15 years ; meaning this small center of wheat evolution is now irridicated. However the former selected lines of the Iranian spelta are being maintained in the gene bank at Braunschweig-Volkenrode. Furthermore the findings of Secale vavilovii, a rather rare wild species proved to be of great value for breeding purposes and research in other fields as well. All data on this matter are summarized in Table 4. I succeeded to collect two samples of this species which are located in a distance of 63 and 9 km from Hamadan respectively : one sample as weed in a wheat field and the second on a steep road side.The progenies of both samples turned out to be completely identical and uniform. The genome proved to be very stable as to spontaneous mutations and also after treatment by X-rays. S. vavilovii is characterized by fragile ears, very small grains and cleistogamous flowers. It is fully self fertil. Samples were distributed to about 20 research institutes the results of which have been presented in about 12 publications up to now (KRANZ 1957, 1973, KUCKUCK und KRANZ 1953, KUSH 1963). Genetic-cytological studies carried out with a view to problems of evolution of cultivated rye revealed the fact that Secale vavilovii is differentiated from S. cereale by two translocations and some smaller chromosomal differences. However the chromosome number of both species is the same 2n=14 (KRANZ 1961, 1973, KUSH 1963).



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