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(c) Ae. cylindrica was found in the Tehran region which is supposed to be the eastern limit of its distribution.

(d) Ae. triuncialis included 2 subspecies, ssp. eu-triuncialis and ssp. orientalis. Ssp. eu-triuncialis was found in all parts of Iran, except for the coastal area of the Caspian Sea; it was also found in the Pul-i-Khumri region of Afghanistan. However, ssp. orientalis was found only in the Maimana region.

(e) Ae. columnaris was found in the Tabriz and Tehran regions. This is a new discovery for its geographical distribution.

2) Morphological characters: The morphological characters such as plant height, tillering habit (procumbent or erect), waxiness (waxy or non-waxy) of leaves, colour of seedlings, color of ears, hairiness (pubescent or glabrous) of glumes, and awnedness of glumes, etc. were studied. Wide variations were found especially in Ae. squarrosa and Ae. triuncialis. For instance, tall plants with erect culms and large grains were found in Ae. squarrosa strains collected from wheat fields, while variations were slight in Ae. cylindrica.

Ae. crassa with awnless ear was found in the south-eastern province of Afghanistan. This will be a new variety.

3) Physiological characters: Earliness, seed-fertility, winter or spring habit, and also resistance to rusts. Some strains of Ae. squarrosa collected in the south eastern provinces of Afghanistan had spring habit, while all other Ae. squarrosa strains exhibited winter habit.

Some strains of var. meyeri and strangulata of Ae. squarrosa have been found to be resistant to certain rust strains.

4) Hybrids of inter- and intraregional cross-combinations were raised among Ae. squarrosa strains. Fertility of those F1 hybrids was normal in most cases, but it was low in some combinations. The sterility of the intra-species hybrids seemed to depend mainly on genotype difference between parents (due to a complementary gene system for hybrid sterility) and partly on environmental conditions.

5) Based on cytological studies, a reciprocal translocation was ascertained in one strain of Ae. squarrosa and two strains of Ae. columnaris found in Iran.



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