| KIHARA and KONDO (1943) succeeded in synthesizing Ae. triuncialis
as an amphidiploid CCCuCu, from caudata x umbellulada,
which resembled var. typica morphologically. They found also that
the ears of the synthesized one (caudata x umbellulata) disarticulate
in the barrel type like those of Ae. squarrosa and crassa.
Based on the observations on chromosome conjugation and fertility in F1
and F2 hybrids between var. typica, or var. persica
and the synthesized one, they concluded that the genomes of them the two
varieties are identical or almost identical. However, further observations revealed that always one or more multivalents and certain number of univalents occur in almost all PMC's in the hybrid of var. typica as one parent. Ssp. orientalis was collected in the Maimana region in Afghanistan by KUSE (1955), but the majority of the collection by BMUK (1959) were ssp. eu-triunrcialis, and a few specimens collected from Konya-Akseki in south-western Turkey were ssp. orientalis var. assyriaca. Since ssp. orientalis was found only in the Maimana region, it was thought that ssp. orientalis was derived from ssp. eu-triuncialis there. But it was not clear if it was originated by mutation of one or more genes concerning the awning and disarticulation, or if it was the product of introgressive hybridization of eu-triuncialis x crassa as proposed by ZOHARY and FELDMAN (1962). However, it was interesting to know that ssp. orientalis was found from Konya-Akseki in Turkey where it was presumed to be the centre of the distribution of Ae. triuncialis, and where awned form of Ae. caudata var. polyathera, having C-genome, occurred widely. |
| 1) The Kyoto University Scientific Expedition to the Karakoram and Hindukush,
1955 2) The Botanical Mission of the University of Kyoto to the Eastern Mediterranean Countries, 1959 |
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