Spontaneous;Bivoltine as normal allele; univoltine(V1, 1 化性) caused by enhanced production of diapausing hormone (DH); multivoltine(V3, 多化性) caused by reduced production of DH; affected to a great extent by Lm alleles(1-2.0): Uptake of 3-hydroxykynurenine by the ovaries of non-diapausing egg layers are in diffusion fashion instead of active transport by the diapausing egg laying ones.