About GAIN
Results of GAIN
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Toshifumi Nakagawa, Masanori Doi, Kosuke Nishi, Takuya Sugahara(2024) Cross-species amplification from non-human primate DNAs in commercial human DNA assay kits
*Legal Medicine*, vol68, Article 102369
https://doi.org/10.1016/j.legalmed.2023.102369
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Tomonaga, M., Tanaka, Y. & Sakai, M.(2024) May the force be with you: exploring force discrimination in chimpanzees using the force-feedback device
*Primates*
https://doi.org/10.1007/s10329-023-01117-1
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Emura K, Hirasaki E, Arakawa T(2023) Muscle–tendon arrangement and intramuscular nerve distribution of flexor digitorum superficialis in the siamang (𝑆𝑦𝑚𝑝ℎ𝑎𝑙𝑎𝑛𝑔𝑢𝑠 𝑠𝑦𝑛𝑑𝑎𝑐𝑡𝑦𝑙𝑢𝑠), western lowland gorilla (𝐺𝑜𝑟𝑖𝑙𝑙𝑎 𝑔𝑜𝑟𝑖𝑙𝑙𝑎 𝑔𝑜𝑟𝑖𝑙𝑙𝑎), western chimpanzee (𝑃𝑎𝑛 𝑡𝑟𝑜𝑔𝑙𝑜𝑑𝑦𝑡𝑒𝑠 𝑣𝑒𝑟𝑢𝑠), and Japanese macaque (𝑀𝑎𝑐𝑎𝑐𝑎 𝑓𝑢𝑠𝑐𝑎𝑡𝑎).
*Anatomical Science International*, in press
https://doi.org/10.1007/s12565-023-00713-x
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Masaki Tomonaga, Yoshiki Kurosawa, Yuri Kawaguchi & Hiroya Takiyama(2023) Don't look back on failure: spontaneous uncertainty monitoring in chimpanzees
*Learning & Behavior*
https://doi.org/10.3758/s13420-023-00581-5
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Masaki Tomonaga & Fumito Kawakami(2023) Do chimpanzees see a face on Mars? A search for face pareidolia in chimpanzees
*Animal Cognition*, 26, 885–905
https://doi.org/10.1007/s10071-022-01739-w
Published: 30 December 2022
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Masaki Tomonaga & Tomoko Imura(2023) Going forward: perceptual bias for forward-facing motion in chimpanzees
*Primates*, 64, 47–63
https://doi.org/10.1007/s10329-022-01034-9
Published: 24 November 2022
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Risako Nakai, Yusuke Hamazaki, Haruka Ito, Masanori Imamura(2022) Early neurogenic properties of iPSC-derived neurosphere formation in Japanese macaque monkeys
*Differentiation*, Vol 128, 33-42.
https://doi.org/10.1016/j.diff.2022.10.002
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Mayu Hirata, Tomoko Ichiyanagi, Hirokazu Katoh, Takuma Hashimoto, Hikaru Suzuki, Hirohisa Nitta, Masaki Kawase, Risako Nakai, Masanori Imamura, Kenji Ichiyanagi(2022) Sequence Divergence and Retrotransposon Insertion Underlie Interspecific Epigenetic Differences in Primates
*Molecular Biology and Evolution*, 39 (10), msac208.
https://doi.org/10.1093/molbev/msac208
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Sakuraya T, Sekiya S, Emura K, Sonomura T, Hirasaki E, Arakawa T(2022) Comparison of the soleus and plantaris muscles in humans and other primates: Macroscopic neuromuscular anatomy and evolutionary significance
*The Anatomical Record*, 306 (2), 386-400.
https://doi.org/10.1002/ar.24999
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Kawada M., Nakatsukasa M., Nishimura T., Kaneko A., Ogihara N., Yamada S., Coudyzer W., Zollikofer C. P. E., De Leon M. S. P., Morimoto N(2022) Human shoulder development is adapted to obstetrical constraints.
*Proceedings of the National Academy of Sciences of the United States of America*, 119 (16) e2114935119
https://doi.org/10.1073/pnas.2114935119
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Takeshi Nishimura, Isao T. Tokuda, Shigehiro Miyachi, Jacob C. Dunn, Christian T. Herbst, Kazuyoshi Ishimura, Akihisa Kaneko, Yuki Kinoshita, Hiroki Koda, Jaap P. P. Saers, Hirohiko Imai, Tetsuya Matsuda, Ole Næsbye Larsen, Uwe Jürgens, Hideki Hirabayashi, Shozo Kojima, W. Tecumseh Fitch(2022) Evolutionary loss of complexity in human vocal anatomy as an adaptation for speech
*Science*, Vol 377, Issue 6607, pp. 760-763
https://doi.org/10.1126/science.abm1574
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Yutaro Sato, Fumihiro Kano, Naruki Morimura, Masaki Tomonaga, and Satoshi Hirata(2022) Chimpanzees (𝘗𝘢𝘯 𝘵𝘳𝘰𝘨𝘭𝘰𝘥𝘺𝘵𝘦𝘴) exhibit gaze bias for snakes upon hearing alarm calls
*Journal of Comparative Psychology*, 136(1), 44–53
https://doi.org/10.1037/com0000305
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Daisuke Kubo(2022) Internal carotid supply to the parietal meninges: a comparative study based on cranio-orbital vascular traces in modern humans and Indonesian 𝘏𝘰𝘮𝘰 𝘦𝘳𝘦𝘤𝘵𝘶𝘴
*Anthropological Science*, 130 (2), 107-119
https://doi.org/10.1537/ase.211223
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Kohta Ito, Tomoya Nakamura, Ryo Suzuki, Takuo Negishi, Motoharu Oishi, Takeo Nagura, Masahiro Jinzaki, and Naomichi Ogihara(2022) Comparative functional morphology of human and chimpanzee feet based on three-dimensional finite element analysis
*Frontiers in Bioengineering and Biotechnology*, 9: 760486
https://doi.org/10.3389/fbioe.2021.760486
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T. Miyabe-Nishiwaki, A. Kaneko, A. Yamanaka, N. Maeda, J. Suzuki, M. Tomonaga, T. Matsuzawa, K. Muta, R. Nishimura, I. Yajima, D. J. Eleveld, A. R. Absalom & K. Masui(2021) Propofol infusions using a human target controlled infusion (TCI) pump in chimpanzees (𝘗𝘢𝘯 𝘵𝘳𝘰𝘨𝘭𝘰𝘥𝘺𝘵𝘦𝘴)
*Scientific Reports*, 11, 1214
https://doi.org/10.1038/s41598-020-79914-7
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Judy Che-Castaldo, Kristin Havercamp, Koshiro Watanuki, Tetsuro Matsuzawa, Satoshi Hirata, Stephen R. Ross(2021) Comparative survival analyses among captive chimpanzees (𝘗𝘢𝘯 𝘵𝘳𝘰𝘨𝘭𝘰𝘥𝘺𝘵𝘦𝘴) in America and Japan
*PeerJ*, 9:e11913
https://doi.org/10.7717/peerj.11913
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Yasuhiro Kikuchi & Naomichi Ogihara(2021) Functional anatomy and adaptation of the third to sixth thoracic vertebrae in primates using three-dimensional geometric morphometrics
*Primates*, 62, 845–855
https://doi.org/10.1007/s10329-021-00929-3
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Takuo Negishi, Kohta Ito, Koh Hosoda, Takeo Nagura, Tomohiko Ota, Nobuaki Imanishi, Masahiro Jinzaki, Motoharu Oishi and Naomichi Ogihara(2021) Comparative radiographic analysis of three-dimensional innate mobility of the foot bones under axial loading of humans and African great apes
*Royal Society Open Science*, 8: 211344
https://doi.org/10.1098/rsos.211344
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Shuhei Nozaki, Hideki Amano, Motoharu Oishi & Naomichi Ogihara(2021) Morphological differences in the calcaneus among extant great apes investigated by three-dimensional geometric morphometrics
*Scientific Reports*, 11, 20889
https://doi.org/10.1038/s41598-021-99942-1
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Shuhei Nozaki, Motoharu Oishi & Naomichi Ogihara(2021) Talar trochlear morphology may not be a good skeletal indicator of locomotor behavior in humans and great apes
*Scientific Reports*, 11, 24063
https://doi.org/10.1038/s41598-021-03429-y
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Susumu Iwaide, Tokuma Yanai & Tomoaki Murakami(2021) Proteopathic lesions in the brain of a super-aged chimpanzee (Pan troglodytes)
*Journal of Medical Primatology*, Vol. 50, Issue4, 222-224.
https://doi.org/10.1111/jmp.12530
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Kinoshita, K., Indo, Y., Tajima, T., Kuze, N., Miyakawa, E., Kobayashi, T., Nakamura, T., Ogata, M., Okumura, F., Hayakawa, T., Morimura, N., Mori, Y., Okamoto, M., Ozaki, Y. & Hirata, S.(2021) Comparative analysis of sperm motility in liquid and seminal coagulum portions between Bornean orangutan (Pongo pygmaeus) and chimpanzee (Pan troglodytes)
*Primates*, 62: 467-473
https://doi.org/10.1007/s10329-020-00887-2
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Brooks, J., Onishi, E., Clark, I., Bohm, M. & Yamamoto, S.(2021) Uniting against a common enemy: perceived outgroup threat elicits ingroup cohesion in chimpanzees
*PLoS ONE*, 16(2): e0246869
https://doi.org/10.1371/journal.pone.0246869
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Brooks, J., Kano, F., Sato, Y., Yeow, H., Morimura, N., Nagasawa, M., Kikusui, T. & Yamamoto, S.(2021) Divergent effects of oxytocin on eye contact in bonobos and chimpanzees
*Psychoneuroendocrinology*, 125:105119
https://doi.org/10.1016/j.psyneuen.2020.105119
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Hee-Eun Lee, Sang-Je Park, Jae-Won Huh, Hiroo Imai & Heui-Soo Kim(2021) The enhancer activity of long interspersed nuclear element derived microRNA 625 induced by NF-κB
*Scientific Reports*, 11, Article number: 3139
https://doi.org/10.1038/s41598-021-82735-x
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Yasuka Toda, Takashi Hayakawa, Akihiro Itoigawa, Yosuke Kurihara, Tomoya Nakagita, Masahiro Hayashi, Ryuichi Ashino, Amanda D. Melin, Yoshiro Ishimaru, Shoji Kawamura, Hiroo Imai & Takumi Misaka(2021) Evolution of the primate glutamate taste sensor from a nucleotide sensor
*Current Biology*, Volume 31, Issue 20
https://doi.org/10.1016/j.cub.2021.08.002
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Jie Gao, Fumito Kawakami & Masaki Tomonaga(2020) Body perception in chimpanzees and humans: The expert effect
*Scientific Reports *, 10, 7148
https://doi.org/10.1038/s41598-020-63876-x
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Torres-Tamayo N, Martelli S, Garcia-Martinez D, Schlager S, Sanchis-Gimeno JA, Nalla S, Ogihara N, Oishi M, Bastir M(2020) Assessing thoraco-pelvic covariation in Homo sapiens and Pan troglodytes: A 3D geometric morphometric approach
*American Journal of Physical Anthropology*, 173(3), 514-534
https://doi.org/10.1002/ajpa.24103
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Ryunosuke Kitajima, Risako Nakai, Takuya Imamura, Tomonori Kameda, Daiki Kozuka, Hirohisa Hirai, Haruka Ito, Hiroo Imai, Masanori Imamura(2020) Modeling of early neural development in vitro by direct neurosphere formation culture of chimpanzee induced pluripotent stem cells
*Stem Cell Research*, 44:101749
https://doi.org/10.1016/j.scr.2020.101749
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Zachary Yu-Ching Lin, Risako Nakai, Hirohisa Hirai, Daiki Kozuka, Seiya Katayama, Shin-ichiro Nakamura, Sawako Okada, Ryunosuke Kitajima, Hiroo Imai, Hideyuki Okano, Masanori Imamura(2020) Reprogramming of chimpanzee fibroblasts into a multipotent cancerous but not fully pluripotent state by transducing iPSC factors in 2i/LIF culture
*Differentiation*, 112: 67-76
https://doi.org/10.1016/j.diff.2020.01.002
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Nakatsukasa M, Morimoto N, Nishimura T(2019) Sesamoids of the pollical metacarpophalangeal joint and the evolution of hominoid hands
*Anthropological Science*, 127(3), 159-164
https://doi.org/10.1537/ase.190908
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Aaron B. Bogutz, Julie Brind’Amour, Hisato Kobayashi, Kristoffer N. Jensen, Kazuhiko Nakabayashi, Hiroo Imai, Matthew C. Lorincz & Louis Lefebvre(2019) Evolution of imprinting via lineage-specific insertion of retroviral promoters
*Nature communications*, 10, 5674
https://doi.org/10.1038/s41467-019-13662-9
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Kristin Havercamp, Koshiro Watanuki, Masaki Tomonaga, Tetsuro Matsuzawa, Satoshi Hirata(2019) Longevity and mortality of captive chimpanzees in Japan from 1921 to 2018
*Primates*, 60: 525-535
https://doi.org/10.1007/s10329-019-00755-8
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Matsuo T, Morita F, Tani D, Nakamura H, Higurashi Y, Ohgi J, Luziga C, Wada N(2019) Anatomical variation of habitat‐related changes in scapular morphology
*Anatomia Histologia Embryologia*, 48(3):218-227
https://doi.org/10.1111/ahe.12426
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Nami Arakawa, Daisuke Utsumi, Kenzo Takahashi, Akiko Matsumoto-Oda, Atunga Nyachieo, Daniel Chai, Ngalla Jillani, Hiroo Imai, Yoko Satta, Yohey Terai(2019) Expression changes of structural protein genes may be related to adaptive skin characteristics specific to humans
*Genome Biology and Evolution*, 11(3): 613-628
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Sakai T, Hata J, Ohta H, Shintaku Y, Kimura N, Ogawa Y, Sogabe K, Mori S, Okano HJ, Hamada Y, Shibata S, Okano H, Oishi K(2018) The Japan Monkey Centre Primates Brain Imaging Repository for comparative neuroscience: an archive of digital records including records for endangered species
*Primates*, 59(6): 553-570
https://doi.org/10.1007/s10329-018-0694-3
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Miki Kakinuma, Hitoshi Hatakeyama, Asami Tsuchida, Izuru Nose, Miki Kamata, and Yukiho Kotani(2018) How would gross motor development leads to changes the quality of mother-child relationship in captive chimpanzees (Pan troglodytes)
*Bull. Nippon Vet. Life Sci. Univ.*, 67, 11-17
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Xu C, Li Q, Efimova O, He L, Tatsumoto S, Stepanova V, Oishi T, Udono T, Yamaguchi K, Shigenobu S, Kakita A, Nawa H, Khaitovich P, Go Y(2018) Human-specific features of spatial gene expression and regulation in eight brain regions
*Genome Res*, 28: 1097-1110
https://doi.org/10.1101/gr.231357.117
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Hirasaki E, Oishi M(2018) Arrangement of foot interosseous muscles in African great apes
*American Journal of Physical Anthropology*, 167, p924-929
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Oishi M, Ogihara N, Shimizu D, Kikuchi Y, Endo H, Une Y, Soeta S, Amasaki H, Ichihara N(2018) Multivariate analysis of variations in intrinsic foot musculature among hominoids
*Journal of Anatomy*, 232, p812-823
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Naoki Morimoto, Masato Nakatsukasa, Marcia S. Ponce de León & Christoph P. E. Zollikofer(2018) Femoral ontogeny in humans and great apes and its implications for their last common ancestor
*Scientific Reports*, volume 8, Article number: 1930
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Hirata S, Hirai H, Nogami E, Morimura N and Udono T.(2017) Chimpanzee Down syndrome: a case study of trisomy 22 in a captive chimpanzee
*Primates*, 58(2), 267-273
https://doi.org/10.1007/s10329-017-0597-8
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Uesaka M, Agata K, Oishi T, Nakashima K, Imamura T(2017) Evolutionary acquisition of promoter-associated non-coding RNA (pancRNA) repertoires diversifies species-dependent gene activation mechanisms in mammals
*BMC Genomics*, 18: 285
https://doi.org/10.1186/s12864-017-3662-1
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Bastir M, García-Martínez D, Williams SA, Recheis W, Torres-Sáncheze I, Río FG, Oishi M, Ogihara N(2017) 3D geometric morphometrics of thorax variation and allometry in Hominoidea
*Journal of Human Evolution*, 113, p10-23
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Shoji Tatsumoto, Yasuhiro Go, Kentaro Fukuta, Hideki Noguchi, Takashi Hayakawa, Masaki Tomonaga, Hirohisa Hirai, Tetsuro Matsuzawa, Kiyokazu Agata & Asao Fujiyama(2017) Direct estimation of de novo mutation rates in a chimpanzee parent-offspring trio by ultra-deep whole genome sequencing.
*Scientific Reports*, 7, 13561.
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Luca Morino, Makiko Uchikoshi, Fred Bercovitch, William D Hopkins, Tetsuro Matsuzawa(2017) Tube task hand preference in captive hylobatids,
*Primates*, 58(3), 403-412
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Sato-Akuhara N, Horio N, Kato-Namba A, Yoshikawa K, Niimura Y, Ihara S, Shirasu M, Touhara K(2016) Ligand specificity and evolution of mammalian musk odor receptors: the effect of single receptor deletion on odor detection
*Journal of Neuroscience*, 36 (16) 4482-4491
https://doi.org/10.1523/JNEUROSCI.3259-15.2016
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Miki Kakinuma(2016) Development of captive chimpanzees at Tama Zoological Park: 15 years of observation with focus on mother-infant relationship
*Japanese Journal of Animal Psychology*, 66(1), 45
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Masahito Irie, Akihiko Koga, Tomoko Kaneko-Ishino, and FumitoshiIshino(2016) An LTR Retrotransposon- Derived Gene Displays Lineage-Specific Structural and Putative Species-Specific Functional Variations in Eutherians
*Frontiers in Chemistry*, 4:26.
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Francesco Nicola Carelli, Takashi Hayakawa, Yasuhiro Go, Hiroo Imai, Maria Warnefors, Henrik Kaessmann(2016) The life history of retrocopies illuminates the evolution of new mammalian genes.
*Genome Research*, 26(3), 301-314.
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Miki Kakinuma, Hitoshi Hatakeyama, Asami Tsuchida, Izuru Nose(2015) Diversities in the Development of Social, Physical and Cognitive Abilities of Captive Chimpanzees (Pan Troglodytes) Compared
*Bull. Nippon Vet. Life Sci. Univ.*, 64,13-19
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Naoki Morimoto, Gen Suwa, Takeshi Nishimura, Marcia S. Ponce de León, Christoph P. E. Zollikofer, C. Owen Lovejoy and Masato Nakatsukasa(2015) Let bone and muscle talk together: a study of real and virtual dissection and its implications for femoral musculoskeletal structure of chimpanzees
*Journal of Anatomy*, 226(3), 258-267.
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Yasumitsu Fujie, Noemi Fusaki, Tomohiko Katayama, Makoto Hamasaki, Yumi Soejima, Minami Soga, Hiroshi Ban, Mamoru Hasegawa, Satoshi Yamashita, Shigemi Kimura, Saori Suzuki, Tetsuro Matsuzawa, Hirofumi Akari, Takumi Era(2014) New Type of Sendai Virus Vector Provides Transgene-Free iPS Cells Derived from Chimpanzee Blood.
*PLOS ONE*, 9(12): e113052
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K. Watanuki, T. Ochiai, S. Hirata, N. Morimura, M. Tomonaga, G. Idani and T. Matsuzawa(2014) Review and Long-Term Survey of the Status of Captive Chimpanzees in Japan in 1926-2013
*Primate Research*, 30(1), 147-156
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Masahiro Uesaka, Osamu Nishimura, Yasuhiro Go, Kinichi Nakashima, Kiyokazu Agata and Takuya Imamura(2014) Bidirectional promoters are the major source of gene activation-associated non-coding RNAs in mammals
*BMC Genomics*, 15:35
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Miki Kakinuma, Hitoshi Hatakeyama, Asami Tsuchida, Izuru Nose(2013) The development of a nursery reared chimpanzee( Pan troglodytes) after joining a socially complex group.
*Bull. Nippon Vet. Life Sci. Univ.*, 62, 68-75.
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Suwa, G., Lovejoy, C. O., Asfaw, B. and White, T.(2012) Proximal Femoral Musculoskeletal Morphology of Chimpanzees and its Evolutionary Significance: A Critique of Morimoto et al. (2011).
*Anatomical Record*, 295, 2039-2044.
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T. Hayakawa, T. Sugawara, Y. Go, T. Udono, H. Hirai, and H. Imai(2012) Eco-Geographical Diversification of Bitter Taste Receptor Genes (TAS2Rs) among Subspecies of Chimpanzees (Pan troglodytes)
*PLOS ONE*, 7(8): e43277.
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N. Morimoto, MS. P. de Leon, T. Nishimura, and CPE. Zollikofer(2011) Femoral Morphology and Femoropelvic Musculoskeletal Anatomy of Humans and Great Apes: A Comparative Virtopsy Study.
*Anatomical Record*, 294, 1433-1445.
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K. Iwamoto, M. Bundo, J. Ueda,M. C. Oldham, W. Ukai, E. Hashimoto,T. Saito,D. H. Geschwind, and T. Kato(2011) Neurons show distinctive DNA methylation profile and higher interindividual variations compared with non-neurons.
*Genome Research*, 21, 688-96.
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T. Sugawara, Y. Go, T. Udono, N. Morimura, M. Tomonaga, H. Hirai and H. Imai(2011) Diversification of bitter taste receptor gene family in western chimpanzees.
*Molecular Biology and Evolution*, 28, 921-931.
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Y. Sato, M.Tomonaga(2010) WAZA (World Association of Zoos and Aquariums) Ethical Guidelines for the Conduct of Research on Animals by Zoos and Aquariums.
*Animal Psychology*, 60, 139-146.
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N. Morimura, G. Idani, and T. Matsuzawa(2010) The first chimpanzee sanctuary in Japan: an attempt to care for the "surplus" of biomedical research.
*American Journal of Primatology*, 71:1-7.
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K. Ohshima, and K. Igarashi(2010) Inference for the initial stage of domain shuffling: tracing the evolutionary fate of the PIPSL retrogene in hominoids.
*Molecular Biology and Evolution*, 27, 2522-2533.
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Alvergne A., Huchard, E. Caillaud, D. Charpentier, M.J.E. Setchell, J. M. Ruppli, C. Fejan, D. Martinez, L. Cowlishaw, G. and Raymond, M.,(2009) Human Ability to Recognize Kin Visually Within Primates.
*International Journal of Primatology*, 30, 199-210.
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T. Ochiai-Ohira(2009) The Kinds of Organization as Facilities Keeping Great Apes in Japan.
*Primate Research*, 24, 405-411.
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M. Oishi, N. Ogihara, H. Endo and M. Asari(2008) Muscle Architecture of the Upper Limb in the Orangutan.
*Primates*, 49: 204-209.
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T. Kawashima, S. Yoshitomi, and H. Sasaki(2007) Nerve Fibre Tracing of Branches to the Coracobrachialis Muscle in a Bornean Orangutan (*Pongo pygmaeus pygmaeus*).
*Anatomia, Histologia, Embryologia*, 36: 19-23.
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M. Kakinuma, H. Hatakeyama, A. Tsuchida and K. Uemura(2007) Difficulties chimpanzee mothers under captivities face in providing maternal care : early maternal separation or social deprivation are not the only reason [in Japanese]
*The bulletin of the Nippon Veterinary and Life Science University*, 56, 28-35.
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Bak,E.J., Ishii,Y., Omatsu,T., Kyuwa,S., Tanoue,T., Hayasaka,I., Yoshikawa,Y.(2006b) Sequence Analysis of Major Histocompatibility Complex Class-II DQB1 (Patr-DQB1) Alleles in Chimpanzees by Polymerase Chain Reaction-Based Methods.
*Human Immunology*, 67, 655-663.
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Bak,E.J., Ishii,Y., Omatsu,T., Kyuwa,S., Tanoue,T., Hayasaka,I., Yoshikawa,Y.(2006a) Identification and analysis of MHC class II DRB1 (Patr-DRB1) alleles in chimpanzees.
*Tissue Antigens*, 67, 134-142.
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Y. Uno, H. Fujino, G. Kito, T. Kamataki, and R. Nagata(2006) CYP2C76, a Novel Cytochrome P450 in Cynomolgus Monkey, is a Major CYP2C in Liver, Metabolizing Tolbutamide and Testosterone.
*Molecular Pharmacology*, 70, 477-486.
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T. Kitano, W.Tian, K. Umetsu, I. Yuasa, K. Yamazaki, N. Saitou, and M. Osawa(2006) Origin and Evolution of Gene for Prolactin-induced Protein,
*Gene*, 383, 64-70.
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T. Ochiai-Ohira, O. Kurashima, R. Akami, T. Hasegawa, M. Hirai, T. Matsuzawa and Y. Yoshikawa(2006) Past and Present of Great Apes in Japan.
*Primate Research*, 22, 123-136.